I watched an SNL skit called "The Californians" on both Hulu and Youtube to see if I could pick out some differences between the two. Hulu, immediately, seemed to jump out as the more "official" version of the two. It seems readily endorsed by the companies behind the film, and not exactly user-submitted. This differs greatly from the Youtube version, which was actually a person's shaky recording of the show that he/she was watching on television at the time. I wonder if this violates Youtube's copyright infringement rules--the video was added 2 days ago so it's relatively new.

As well, Youtube has the percentage of likes/dislikes available in search results, whereas you have to click into the "reviews" section of a Hulu video in order to have an idea of whether people liked the video or not. Hulu also separates "Reviews" and "Discussions," while in Youtube one combines the two (and random pieces of spam) in the comments section.
The audiences for Youtube and Hulu differ as well. Youtube audiences are more casual, and probably younger, as evidenced by the tone of the comments. An example of a Youtube comment on the video is "i laughed so damn hard during this skit!!" while on Hulu a typical review is "As a Californian, I have no idea what is going on in this...but it cracked me up! The references to freeways is pretty dead on. The character breaks were awesome."
There was a slight difference in ways one could share the video--Youtube and Hulu both have links to Twitter and Facebook, but Youtube has a link to Google+ as well. If you click onto other areas of the site on Hulu, you have to pay for Hulu Plus in order to access certain parts of the site or certain videos that have been aired a longer time ago.
In summary, Hulu is more "corporate," and Youtube is more "user-produced."
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